This story picks up a generation after the famous race between the Hare and the Tortoise. This time it's their kids' turn to compete, and they're about to escalate their family rivalry to ridiculous new levels.

After losing everything, a young fish, Pi, goes to live with his family on the Reef. There he meets the love of his life but finds that she already has the unwanted affections of a bully shark. He must follow his destiny to save her and rid the Reef of this menace for good.